By Kennedy W.J., Cobban W.A.
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Extra resources for Aspects of ammonite biology, biogeography and biosratigraphy
Flexuosum (of which A. crassiornatum Crick, A. expansum Crick, A. robustum Crick, A. quadratum Crick, A. hippocastanum Crick (non Sower by), and A. latum Crick are probably synonyms). In Texas a rather similar group of ammonites is represented by the AMMONITE ECOLOGY 41 A. stephensoni Adkins, A. adkinsi Stephenson-v4. , tarrantense Stephenson complex. Individuals from any one population would pass unnoticed in any other, but over-all population structures are clearly different. All faunas are of Middle Ceno manian age, but correlation of these widely separated areas is too imprecise to determine their relative ages accurately.
Washings of sediments deposited many miles from shore often yield abundant juvenile ammonites (A. J. Lloyd, pers. ), and we know of many cases of huge concentrations of juveniles, complete with body chambers, preserved in pelagic deposits. For example we have a collection from the Marias River Shale (Santonian) of north-western Montana, which is a mass of juvenile Baculites codyensis Reeside and Clioscaphites vermiformis (Meek and Hayden). These specimens are complete with body chambers and aper tures; they died possibly 150 km east of the western shoreline of the Western Interior epeiric sea.
Nine species have been attributed to the genus, but unpublished work by Kennedy and Klinger suggests that only three appear to merit separation as distinct morphospecies. These species have overlapping stratigraphical and geographical ranges, and may represent no more than variants or subspecies of a P. indra (Forbes) biospecies. Pseudophyllites is common in the Campanian/Maastrichtian of Alaska, the then North Pole (Smith et al. 1973), British Columbia, and East Greenland. It is also common in Antarctica (Graham Land), South Africa, Madagascar, Australia, and New Zealand, at that time in high southern latitudes.